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U.S. authorities said they had broken up a plot by two men linked to Iran's security agencies to assassinate Saudi Ambassador Adel al-Jubeir. One was arrested last month while the other was believed to be in Iran.
Iran denied the charges. But President Barack Obama called the plot a "flagrant violation of U.S. and international law" and Saudi Arabia said it was "despicable."
Gen Bergner said Mr Dakdouk had told his US interrogators that the Karbala attackers "could not have conducted this complex operation without the support and direction of the Quds force".
Five Americans were killed in the Karbala assault, a bold and sophisticated assault in which up to a dozen gunmen posed as a US security team to gain access to the government compound.