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Lonesome George, the last remaining tortoise of his kind and a conservation icon, has died of unknown causes in Ecuador's Galapagos Islands.
The giant tortoise was found in 1972, and was thought to be about 100 years old.
Lonesome George was a symbol of Ecuador's Galapagos Islands, which attracted 180,000 visitors last year.
The head of the Galapagos National Park, Edwin Naula, says "his life cycle came to an end" on Sunday (local time).
"This morning the park ranger in charge of looking aft
Harriet (c. 1830 - June 23, 2006) was a Galápagos tortoise (Geochelone nigra porteri) who had an estimated age of 175 years at the time of her death in Australia. Harriet is the third oldest tortoise ever authenticated, behind Tu'i Malila, who died in 1965 at the age of 188, and Adwaita, who died in 2006 at the estimated age of 255.