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In 1917 and 1918, close to 5 million Americans served in World War I, and Mr. Buckles, a cordial fellow of gentle humor, was the last known survivor. "I knew there'd be only one someday," he said a few years back. "I didn't think it would be me."
His daughter, Susannah Buckles Flanagan, said Mr. Buckles, a widower, died of natural causes on his West Virginia farm, where she had been caring for him.
When you think you're dying, his son-in-law once heard him quip, don't.