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* Anatomical and behavioural adaptations that allow them to move effectively on land as well as in the water.
* The ability to breathe through their skin and the lining of their mouth (the mucosa) and throat (the pharynx). This is only possible when the mudskipper is wet, limiting mudskippers to humid habitats and requiring that they keep themselves moist. This mode of breathing, similar to that employed by amphibians, is known as cutaneous air breathing. Another important adaptation that aids breathing are their enlarged gill chambers, where they retain air. These act like a scuba diver's oxygen cylinders, and supply oxygen for respiration also while on land.
* Digging of deep burrows in soft sediments that allow the fish to thermoregulate; avoid marine predators during the high tide when the fish and burrow are submerged; and for laying their eggs. Even when their burrow is submerged, mudskippers maintain an air pocket inside it, which allows them to breathe in conditions of very low oxygen concentration.
Pygopodidae is the family of legless lizards. They are distinguished from snakes by their eyelids that can blink (snakes have no eyelids), external ear holes (snakes have no ears at all, internal or external), and flat, non-forked tongues. Many species also feature vestigial limbs, in the form of scaly flaps.
The misconception about the lack of transitional fossils is perpetuated in part by a common way of thinking about categories. When people think about a category like "dog" or "ant," they often subconsciously believe that there is a well-defined boundary around the category, or that there is some eternal ideal form (for philosophers, the Platonic idea) which defines the category. This kind of thinking leads people to declare that Archaeopteryx is "100% bird," when it is clearly a mix of bird and reptile features (with more reptile than bird features, in fact). In truth, categories are man-made and artificial. Nature is not constrained to follow them, and it doesn't.